These are freshwater mussels. 

These animals are freshwater mussels. 

Mussels have a tiny foot. They use it to move. They also use it to bury themselves. This helps them stay safe in the mud. 
Mussels have a special way to grow. They start as tiny babies. These babies must hitch a ride on a fish. They stick to the fish for a while. Then they fall off to grow. This helps them move to new places. They are very good at living in fresh water.
Unionida are a group of freshwater mussels. 

Mussels have a muscular foot. They use this foot to move slowly. They also use it to bury themselves in mud. To eat, they pump water through their bodies. This lets them get oxygen and food from the water.
These mussels have a very strange life cycle. They start as tiny larvae called glochidia. 
People once caught many mussels to make buttons. They also looked for pearls. Because of this, many species are now in trouble. Some are endangered or even extinct. 
Unionida are a special group of living things called freshwater mussels. 

To live, these mussels use a clever way of working. They use a muscular foot to move slowly through the mud. They bury themselves in the bottom of the river. To eat, they pump water through their bodies. This helps them get oxygen and tiny bits of food. The shells have a shiny, pearly inside called nacre. 
The way these mussels grow is very unusual. They start life as tiny larvae called glochidia. 
Humans have a long history with these animals. In the mid-1800s, people went through a "pearl rush." 

Today, many mussels face hard jobs to stay alive. Because so many were taken for buttons and pearls, many species are in trouble. In North America, about 70% of Unionida are extinct, endangered, or threatened.
Unionida is a monophyletic order of aquatic bivalve molluscs known as freshwater mussels. 

The physical structure of these mussels allows them to survive in specific environments. Their shells are typically elongate and equivalve, meaning the two valves are of roughly equal size. The interior features a pearly nacreous layer, radial sculpture, and an entire pallial line. To move, they use a muscular foot to slowly bury themselves in the substrate. They live in clean, fast-flowing freshwater like rivers, streams, and creeks. These mussels pump water through an incurrent aperture to obtain oxygen and filter food. Because their siphons are short, they cannot live deeply buried in sediment. 
Reproduction in Unionida involves a complex and unique life cycle. Most species have separate sexes, though some, like Elliptio complanata, are hermaphroditic. During mating, sperm is ejected from the male's excurrent aperture into the female's incurrent aperture. Fertilized eggs move to the gills, called marsupia, to become glochidia. These are the first larval stage of the mussel. Mature glochidia are released and attach to a host fish using tiny hooks. 
Some species use specialized methods to ensure their larvae find a host. Many release glochidia in mucilaginous packets known as conglutinates. These packets have a sticky filament to prevent them from being washed away. Some mussels produce a super-conglutinate that looks like a fish egg or an aquatic fly larva. This version has a dark eyespot to make it look appetizing to fish. When a fish eats the super-conglutinate, it breaks apart and releases the glochidia. Once attached, the larvae stay on the fish for weeks or months before falling off as juveniles. 
The evolutionary history of Unionida is a subject of scientific study. Some fossils from the Carboniferous and Permian periods were suggested to be unionids. However, many authors disagree because those fossils lack the internal nacreous layer. This layer is found in the last common ancestor of all unionids and their marine relatives, the Trigoniida. The oldest unambiguous unionids come from the Triassic period. These are found in the Chinle Formation and Newark Supergroup of North America. Possible older examples might exist in the Middle Triassic of Tanzania and Zambia.
Humans have significantly impacted Unionida through various commercial industries. In the mid-1800s, a "pearl rush" occurred in North America. People used a method called "pollywogging," where they shuffled their feet in mud to find pearls. This was similar to the California gold rush. Later, in the late 19th and early 20th centuries, the nacre was used for button manufacturing. This heavy exploitation put many species close to extinction. 
Today, many species face serious conservation issues due to habitat degradation and past over-exploitation. In North America, about 70% of Unionida are extinct, endangered, threatened, or of special concern. This includes 21 extinct species and 77 endangered species. Many species, such as the butterfly, mapleleaf, and washboard, were historically harvested. While natural pearls are often irregular, shaped like slugs or wings, the Japanese industry developed pearl culturing. This industry uses nacre from exported North American shells to create bead nuclei for round pearls. 
Despite these challenges, the Unionida remain a vital part of freshwater ecosystems. They are found on six continents, with approximately 900 species worldwide. About 300 of these species are endemic to North America. Their ability to move upstream by hitchhiking on fish is a remarkable survival strategy. Understanding their life cycles and history helps us protect the rivers they inhabit. Many states, including Alabama and Tennessee, remain important areas for mussel production and study.
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